Berteroa physocarpa

Herbs, perennial, moderately pubescent with appressed stellate trichomes. Stems 40–75 cm, ascending, terete, branched at base, leafy, pubescent with 5–7 unequal-rayed stellate trichomes longest rays of which parallel to stem long axis, some appearing as submalpighiaceous. Basal leaves rosulate, oblanceolate, 4–12 × 1–4 cm including basally flattened petioles 1.5–3 cm, pubescent with appressed, sessile stellate trichomes some with central subsetose ray and others submalpighiaceous along entire to coarsely dentate margin, base cuneate to attenuate; middle cauline leaves oblanceolate, lanceolate, to elliptic, 4.5–7 × 1–1.5 cm, sessile, cuneate at base, gradually reduced in size upwards and uppermost to 10 × 2.6 mm. Raceme densely flowered, only slightly elongated in fruit; rachis pubescent as stem; fruiting pedicels 6.5–9.4 mm, ascending to divaricate-ascending, straight or slightly curved, pubescent with stellate trichomes some with reduced rays except subsetose central ray and appearing simple. Sepals 3.7–4.5 mm, ovate-oblong, with membranous white margin, stellate pubescent, subapically with setose central ray and reduced lateral rays, subsaccate at base; petals white, 6.3–8.5 mm, narrowly obcordate, with oblong lobes 2.5–4 mm, attenuate to flattened claw-like base 3–4 mm; median filaments 4.1–4.9 mm, dilated at base, unappendaged; lateral filaments 3.3–4.1 mm, basally with toothed oblong appendage 0.7–0.9 mm; anthers yellow, oblong, 1–1.3 mm; nectar glands 4, toothlike, 1 on each side of lateral stamen; ovules 4–5 per locule. Fruit oblong-ellipsoid to globose, distinctly inflated, 9.3–13.3 × 6.8–8.6 mm, glabrous on valves, occasionally with a few trichomes along proximal part of replum; septum complete, membranous; style slender (2.6–)3–3.8(–4.2) mm, glabrous or with few trichomes at base. Seeds suborbicular-lenticular, strongly flattened, minutely reticulate, 3–4 mm diam., including undulate, usually paler wing 0.4–0.6 mm wide.